List of all taxa, sorted by Subgenus etc

The list of sources used to compile this list is available on the main page for this section.

A version of this list Sorted Alphabetically [120K] is also available.

Section I - Paucispini Fric

Subdivided into the Ottonis Group, Linkii Group and Acutus Group.

Series 1 - Ottonis Group

arechavaletae (Spegazzini) Herter: Cact. Plat. Tentamen, 496-497, Buenos Aires 1905
Flore illustrée l'Uruguay - Cactacées, 177, 1955.
arechavaletae var. alacriportanus Ritter: Kakt. in Süd., 1:165-166, Spangenberg 1979.
arechavaletae var. aureus Ritter: Kakt. in Süd., 1:166-167, Spangenberg 1979.
arechavaletae var. buenekeri Ritter: Kakt. in Süd., 1:166, Spangenberg 1979.
arechavaletae var. convexicostatus n.n.
arechavaletae var. horstii Ritter: Kakt. in Süd., 1:166, Spangenberg 1979.
arechavaletae var. grandicostatus n.n.: Note: Only a name ?
arechavaletae var. limiticola Ritter: Kakt. in Süd., 1:164-165, Spangenberg 1979.
arechavaletae var. nanus Ritter: Kakt. in Süd., 1:165, Spangenberg 1979.
arechavaletae var. rubescens Ritter: Kakt. in Süd., 1:166, Spangenberg 1979.
arechavaletae var. segredoensis n.n.
And varieties from the following localities:: Rivera.
And forms from the following localities:: Bage, Baram Itu, Cruz Alta, Cuchilla Negra, Dom Pedrito, Estancia Platon, Fazenda Santa Victoria, Livramento, Melo, Minas, Pinheiro Machado, Routa 26 / Tacuarembo, Rivera, Tobai and Tranqueras.
batoviensis n.n. [?]
blossfeldianus n.n.: Note: Synonym of ottonis var. blossfeldianus.
campestrensis Ritter: Kakt. in Süd., 1:177, Spangenberg 1979.
electracanthus n.n. [?]
eurypleurus n.n. [?]
glaucinus Ritter: Kakt. in Süd., 1:168, Spangenberg 1979.
glaucinus var. alegretensis n.n.
glaucinus var. depressus Ritter: Kakt. in Süd., 1:169, Spangenberg 1979.
glaucinus var. gracilis Ritter: Kakt. in Süd., 1:168-169, Spangenberg 1979.
Note: Wolfgang Gemmrich considers this to be a distinct species; see Internoto 17 (3):91-99, 1996.
glaucinus var. gracilispinus n.n.
glaucinus var. miniatispinus n.n.
And forms from the following localities:: Cerro de Tigre, Estancia Itu, "Near HU 11", Itaqui, Santiago, S. Santiago and Villa Kramer.
globularis: Note: Reduced to a variety of ottonis.
grandiensis n.n.: Note: Synonym of ottonis var. grandiensis.
harmonianus Ritter: Kakt. in Süd., 1:176-177, Spangenberg 1979.
haasisianus n.n. [?]
ibicuiensis Prestlé: Internoto 6 (4):99-103, 1985.
ibicuiensis var. subtilispinus n.n.
And forms from the following localities:: Estancia Nova.
incomptus Gerloff: Internoto 11 (2-3):37-41, 1990.
intermedius n.n. [?]
laetivirens Ritter: Kakt. in Süd., 1:167-168, Spangenberg 1979.
laetivirens var. tenuispinus n.n.
laetivirens fa. densispinus n.n.
medinaensis n.n. [?]
militaris n.n.: Note: Transitional taxon between Paucispini and Uebelmannianae ?
minusculus Hofacker et Herm [?]: Internoto 17 (3):75-79, 1996
minusculus ssp gravior Hofacker [?]: KuaS, 51(9):233-236, 2000.
Note: Described as Parodia nothominuscula ssp. gravior
muricatus (Otto) Berger ex Backeberg: Enumeratio Diag. Cact., 49, Berlin 1837
Kakteen, 210, Stuttgart 1929.
Note: There has been some confusion over the usage of this name, if your muricatus looks like a member of the Setacei then you probably have bommeljei.
muricatus var. albispinus n.n.: Note: Synonym of muricatus.
muricatus var. monticola n.n.
muricatus var. flavifuscus n.n.
muricatus fa. densispinus Bergner: Internoto 14 (1):3-12, 1993.
muricatus fa. monticola n.n.
muricatus fa. robustus n.n.
nothominuscula Hofacker [?]: Note: The IOS consensus name for minusculus.
oligacanthus n.n. [?]: Note: Provisional name for minusculus.
ottonis (Lehmann) Berger ex Backeberg: Index seminum hort. Hamburg, 16, 1827
Kakteen, 212, Stuttgart 1929
Note: Type species for the Subgenus, Paucispini and Series 1.
ottonis var. brasiliensis n.n.: Note: Synonym of linkii var. brasiliensis.
ottonis var. blossfeldianus n.n. [?]
ottonis var. cacapavensis n.n. [?]
ottonis var. callianthus n.n. [?]
ottonis var. campestrensis n.n. [?]: Note: Synonym of campestrensis.
ottonis var. campoensis n.n. [?]: Note: Synonym of campoensis.
ottonis var. centangularis n.n. [?]
ottonis var. chrysanthus n.n. [?]
ottonis var. densispinus n.n. [?]
ottonis var. denudatus n.n.: Note: Synonym of var. schuldtii.
ottonis var. durispinus n.n. [?]
ottonis var. elegans Backeberg et Voll: Arqu. Jard. Bot. Rio de Janeiro, 9:172, 1949.
ottonis var. globularis (Ritter) Bergner: Kakt. in Süd., 1:167, Spangenberg 1979
Internoto 9 (3):67-73, 1988.
ottonis var. grandiensis n.n.: Prov. Desc.: S. Bergner, Internoto 10 (2):43-47, 1989.
ottonis var. grandiflorus Papousek Reference ???
ottonis var. haseltonianus n.n. [?]: Note: Only a name ?
ottonis var. janousekianus Papousek: Kaktusy 11 (6):123-126, 1975.
ottonis var. knippelianus n.n. [?]: Note: Only a name ?
ottonis var. knesplii n.n. [?]
ottonis var. longispinus n.n. [?]
ottonis var. machadoensis n.n. [?]
ottonis var. mesetaensis n.n. [?]
ottonis var. minasensis n.n. [?]
ottonis var. minimus n.n. [?]: Note: Only a name ?
ottonis var. minor n.n. [?]: Note: Only a name ?
ottonis var. minusculus n.n. [PO ?]: Note: Synonym of minusculus.
ottonis var. mistioensis n.n.: Note: Synonym of fa. Misti.
ottonis var. nigrispinus n.n. [?]: Note: Synonym of var. minasensis
Prov. Desc.: H. D. Lück;, Internoto 8 (4)124-125, 1987.
ottonis var. pantanoensis n.n. [?]
ottonis var. paraguayensis (Heese) Berger ex Backeberg: Gartenwelt, 9:266, 1905
Kakteen, 213, Stuttgart 1929
Note: Original author sometimes mistakenly given as Haage jr.
And forms from the following localities:: Barrero Grande, Capilla Tuya, Carai-mi, Eusebio Ayala, Chololo, Isla Ybate, Itacurubi, Piribebuy, Ruta 2, Serrania de Ybycui (Ybucui ?), Valenzuela, Villa Florida and Yhaguy Guazu.
ottonis var. palitor n.n. [?]: Note: Only a name.
ottonis var. prolifer n.n. [?]
ottonis var. rubrispinus n.n. [?]
ottonis var. rufispinus n.n. [?]
ottonis var. schuldtii Kreuzinger: Succulenta 27 (4):47, 1948.
ottonis var. segredoensis n.n. [?]
ottonis var. spinosior n.n. [?]: Note: Only a name.
ottonis var. tenuispinus (Link et Otto) Berger ex Backeberg: Verh. Ver. Beförd. Gartenb., 3:421, Berlin 1827
Kakteen, 213, Stuttgart 1929.
ottonis var. uruguayus (Arechavaleta) Berger ex Backeberg: Flora Uruguaya, 5:213-215, Montevideo 1905
Kakteen, 214, Stuttgart 1929.
ottonis var. vargasensis n.n. [?]
ottonis var. vencluianus Schütz: Kaktusy 1 (6):123-126, 1965.
And varieties from the following localities:: Piedra de Afilar, Rivera and Treinta-y-Tres [?].
And forms from the following localities:: Abra da Zavaleta / Lavalleja, Acegua, Aigua, Alegreta N, Alegreta S, Arroyo Cunapiru, Arroyo Grande, Artigas, Bage, BR 116/km 225 Sta Catarina, Bura, Cacapava N, Cacapava S, Between Casias and Vacaria, Cerro Areicu, Cerro de Cunapiru, Cerro de los Cuentas, Cerro Grande, Cerro Largo, Cerro Mosquito, Collares, Cordoba, Cuchilla Negra, Cunapiru (Mountain), Curtina, Darilho, Del Torro, Dom Feliciano, Estribo, Estribo de Mineracao, Fronteira, Gruta de los Cuervos, Gruta de Segredo, Gruta del Palacio, Ibare, Incatus, Itaqui, Knespli, Laguna Garzon, Lavras, Lemi, Livramento, Loreto (Mountain), Los Quadros, Maldonado (Loreley), Maldonado, Manuel Vianna, Masoller, Melo, Minas, Minas de Corrales, Morro Embratel, Misti / Uruguay, Neocactel, Nova Palma, Paso del Cerro, Pedras Altas, Pelotas, Pena, Piratini, Piriapolis, Piraraja, Pororo, Porto Alegre, Punta Ballena, Punta de la Arena, Quarai, Rivera, Ruta 30/km 100, San Antonio (Piriapolis), Sao Francisco de Assis, Sao Gabriel, Sao Pedro, Santana do Livramento, Santa Rita, Termas del Daiman, Tacuarembo, Tacuarembo E, Tacuarembo N, Tacuarembo NW, Tacuarembo W, Tigre, Torres, Tranqueras, Treinta-y-Tres, Tupantuba, Uruguaiana and Vichadero [?].
paradoxus n.n. [?]
ruoffii Gerloff: Internoto 14 (4):115-119, 1989
Note: Transitional taxon between Paucispini and Fricianae.
spegazzinii n.n.: Note: Synonym of arechavaletae.

Series 2 - Linkii Group

campoensis n.n.[?]: Prov. Desc.: N. Gerloff, Internoto 6 (3):79-80,1985.
carambeiensis Buining et Brederoo: Kakt. u. a. Sukk. 24 (1):1-3, 1973.
carambeiensis var. villa-velhensis (Buining et Brederoo) Sida ex Dohnke: Kakt. u. a. Sukk. 24 (1):1-3, 1973
Internoto 15 (4):143-144, 1994.
linkii (Lehmann) Herter: Sem. hort. bot. Hamb. 16, 1827
Flore illustrée l'Uruguay - Cactacées, 120, 1955
Note: Type species for the Paucispini - Series 2.
linkii var. albiareolatus Schäfer Reference ???
linkii var. albispinus Abraham: Succulenta 69 (5):117-120, 1990.
linkii var. altosensis n.n.
linkii var. berlinensis n.n.
linkii var. brasiliensis n.n.
linkii var. buenekeri Ritter: Kakt. in Süd., 1:172, Spangenberg 1979.
linkii var. flavispinus: Note: Transferred to a variety of megapotamicus.
linkii var. flavispinus fa. websterianus n.n.: Prov. Desc.: Internoto 9 (2):58-60, 1988.
linkii var. guaibensis Ritter: Kakt. in Süd., 1:172, Spangenberg 1979.
linkii var. jaquiranensis n.n.
linkii var. multiflorus Fric ex Buining sine comb.: Note: Ritter reduced this to a Form (Kakt. in Süd., 1:172, Spangenberg 1979).
linkii var. paranaensis n.n.
linkii var. paucicostatus n.n.
linkii var. stenogonus n.n.
linkii var. tenebrosus n.n.: Prov. Desc.: G. Schäfer and K. Lisal, Internoto 4 (3):61-66, 1983.
linkii var. uruguayensis n.n.: Note: Synonym of megapotamicus fa. S 155.
linkii var. vermelhensis n.n.: Note: Provisional name for linkii var. flavispinus
linkii fa. Itapua: Note: Synonym of megapotamicus var. alacriportanus.
linkii fa. "Ossowski": Note: Synonym of acutus fa. "Ossowski".
And forms from the following localities:: Apostoles, Arroio san Juan, Bom Jardim da Serra, Bom Jesus, Cacapava do Sul, Cachoeira do Sul, Cambara, Cambara "Near HU 42", Casa da Pedro / Bage, Caxias, Cerro Areicu, Curitiba, Fortaleza, Garganta do Diabolo, Itambezinho, Kraporchas, Livramento, Painel Brasil, Quebra Dentas, Rio Grande do Sul, Ruta 5 / km 395, Saltinho, Santa Catarina, Santa Tecla / Bage, Santiago, Sao Gabriel "Near HU 25", Sao Gabriel "Near HU 45", Tahinia and Vacaria "Near HU 39".
megapotamicus (Osten) Herter: Notas sobre Cactáceas, 72-73, 1941
Rev. S. Am. Bot., 7:73-74, 1942.
megapotamicus var. alacriportanus Ritter: Kakt. in Süd., 1:175, Spangenberg 1979.
megapotamicus var. aurea n.n.: Note: Only a name ?
megapotamicus var. catarinensis n.n.: Note: Synonym of megapotamicus fa. Santa Maria.
megapotamicus var. crucicentrus Ritter: Kakt. in Süd., 1:176, Spangenberg 1979.
megapotamicus var. flavispinus (Buining et Brederoo) Abraham and Büneker: Kakt. u. a. Sukk. 29 (2):25-27, 1978
Kakt. u. a. Sukk. 43 (1):20-22, 1992.
megapotamicus var. horstii Ritter: Kakt. in Süd., 1:175-176, Spangenberg 1979.
megapotamicus var. horstii fa. supertextus n.n.
megapotamicus var. multicolorispinus Abraham et Bueneker: Kakt. u. a. Sukk. 43 (1):20-22, 1992.
megapotamicus var. placentiformis n.n.
megapotamicus var. placentiformis fa. supertextus n.n.: Note: Only a name.
megapotamicus var. tenuispinus n.n.
megapotamicus var. vulgatus Ritter: Kakt. in Süd., 1:175, Spangenberg 1979.
megapotamicus fa. multiflorus (Fric ex Buining) Stuchlík: Succulenta 36 (9):101-108, 1957
Rod Notocactus 85-86, 1993.
And forms from the following localities:: Alem de los Cerros, Bage, Cerro de Tigre, Dom Pedrito, Maldonado, Manuel Vianna, Quevedos, Sao Francisco de Assis, Santa Maria, "Near HU 813", Suspiro and Unistalda.
ottonis var. albispinus Backeberg: Note: Synonym of linkii var. albispinus.
ottonis var. brasiliensis (Haage Jr.) Berger: Monatsschrift für Kakteenkunde, 41, 1914
Kakteen, 214, Stuttgart 1929
Note: Synonym of linkii var. brasiliensis.
ottonis var. multiflorus Fric ex Buining: Succulenta 36 (9):101-108, 1957
Note: Transferred to a form of megapotamicus.
ottonis var. nugualensis n.n.: Note: Synonym of var. tortuosus.
ottonis var. pampeanus n.n.: Note: Synonym of var. tortuosus.
ottonis var. stenogonus Backeberg: Die Cactaceae, 6:3758, Jena 1962.
ottonis var. tenebrosus n.n.: Note: Synonym of linkii var. tenebrosus.
ottonis var. tortuosus (Link et Otto) Berger ex Backeberg: Icon. Selt. Gew., 29, 1830
Kakteen, 213, Stuttgart 1929.
ottonis var. villa-velhensis Backeberg et Voll: Note: Synonym of villa-velhensis.
ottonis var. websterianus n.n.: Note: Synonym of linkii var. flavispinus fa. websterianus.
riosusannaensis n.n.: Note: Intermediate between megapotamicus and linkii ?
Prov. Desc.:^K. H. Prestlé, Internoto 6 (1):30-31, 1985.
supertextus n.n.: Note: A form of megapotamicus var. horstii.
villa-velhensis: Note: Reduced to a variety of carambeiensis.

Series 3 - Acutus Group

acutangularis n.n.: Note: Synonym of ottonis var. acutangularis.
acutus: Note: Reduced to a form of oxycostatus.
acutus var. cadeiaensis n.n.: Prov. Desc.: K. H. Prestlé, Internoto 1 (1):2-3, 1981
Note: Synonym of cadeiaensis.
acutus var. lourencoensis n.n.: Prov. Desc.: K. H. Prestlé, Internoto 1 (1):2, 1981.
acutus fa. "Ossowski".
cadeiaensis n.n.
obtectus n.n.: Note: Synonym of securituberculatus ?
ottonis var. acutangularis Ritter: Kakt. in Süd., 1:163, Spangenberg 1979.
oxycostatus Buining et Brederoo: Die Kakteen, C VIc, Stuttgart 15. 10. 1972.
oxycostatus fa. acutus (Ritter) Gerloff: Kakt. in Süd., 1:169-170, Spangenberg 1979
Internoto 14 (3):83-95, 1993.
oxycostatus fa. securituberculatus (Ritter) Gerloff: Kakt. in Süd., 1:169, Spangenberg 1979
Internoto 14 (3):83-95, 1993.
oxycostatus fa. miniatispinus (Ritter) Gerloff: Kakt. in Süd., 1:169, Spangenberg 1979
Internoto 14 (3):83-95, 1993.
paucispinus n.n.: Note: Synonym of acutus.
pseudoacutus n.n. [?]
quevedoi n.n.[PA]: Note: Synonym of acutus.
securituberculatus: Note: Reduced to a form of oxycostatus.
securituberculatus var. miniatispinus: Note: Reduced to a form of oxycostatus.
securituberculatus var. ritteri n.n.

Transitional forms

With Fricianae: ruoffii and stockingeri.

Section II - Fricianae Havlicek

caespitosus (Spegazzini) Backeberg: Note: The name caespitosus has been shown to be a prior name of apricus (Theunissen, Internoto 3 (3):77-81, 1982).
Note: Synonym of minimus.
erectocylindricus n.n.: Note: Synonym of tenuicylindricus PR 250.
minimus Fric et Kreuzinger: Succulenta 22 (8):86-87, 1940
Note: Type species for Section II.
minimus var. albispinus n.n.
minimus var. aureispinus n.n.
minimus var. maximus n.n.
And forms from the following localities:: Alegrete S, Arroyo Cunapiru, Baja de Peguoi, Cuchilla Negra, Dept. Artigas, Masoller, Rivera, Ruta 30 km 45, Ruta 30 km 108, Ruta 5 km 407, Ruta 5 km 412, Salto, St. Ana do Livramento, Tacuarembo and Transqueros.
rengglianus n.n.: Note: Synonym of stockingeri ?
stockingeri Prestlé: Succulenta 64 (11):226-229, 1985
Note: Transitional taxon between Fricianae and Paucispini.
stockingeri var. durispinus n.n.
stockingeri var. semicylindricus n.n. ???
tenuicylindricus Ritter: Succulenta 49 (7):108-109, 1970.
tenuicylindricus var. atypicus n.n.
tenuicylindricus var. caveraensis n.n.

Transitional forms

With Paucispini: ruoffii and stockingeri.

Section III - Scopanae Havlicek

adustus n.n.: Note: Provisional name for soldtianus.
cobrensis n.n.: Note: Synonym of scopa var. cobrensis.
elachisanthus n.n.: Note: Synonym of scopa var. elachisanthus.
fuscus Ritter: Kakt. in Süd., 1:178, Spangenberg 1979
Note: Transitional taxon between Scopanae and Neonotocactus.
fuscus var. longispinus Ritter: Kakt. in Süd., 1:179, Spangenberg 1979.
gerloffianus n.n.
glomeratus Gerloff: Internoto 12 (1):3-10, 1991.
neobuenekeri Ritter: Kakt. in Süd., 1:181-182, Spangenberg 1979.
neobuenekeri fa. caespitosus n.n.
And forms from the following localities:: Santana de Boa Vista.
nothorauschii D. Hunt: Note: The IOS consensus name for rauschii.
pitcayensis n.n.: Note: Var. of scopa ?
rauschii Van Vliet: Succulenta 48 (1):3-5, 1969
Note: Transitional taxon between Scopanae and Neonotocactus.
rauschii var. caveraensis n.n.
And varieties from the following localities:: Rivera / Livramento.
rudibuenekeri Abraham: Succulenta 67 (6):133-138, 1988.
scopa (Sprengel) Berger ex Backeberg: Systema Vegetabilium, 2:494, Göttingen 1825
Kakteen, 208, Stuttgart 1929
Note: Type species for the Section III.
scopa var. adustus n.n.: Note: Provisional name for soldtianus.
scopa var. albicans (Arechavaleta) Hofacker: Flora Uruguaya, 5:199-201, Montevideo 1905
Internoto 11 (2-3):68-74, 1990.
scopa var. albispinus n.n.
scopa var. aureispinus n.n.: Note: Synonym of fa. daenikerianus ?
scopa var. brachyacanthus n.n.: Note: Provisional name for scopa var. cobrensis.
scopa var. brasiliensis n.n.
scopa var. brunneispinus n.n.
scopa var. caputproliferanus n.n.
scopa var. cobrensis Gerloff: Internoto 11 (1):3-9, 1990.
scopa var. caespitosus n.n.: Note: Synonym of fa. candidus or var. ramosus ?
scopa var. elachisanthus n.n.
scopa var. fricii n.n.: Note: Synonym of var. ruberrimus.
scopa var. fuscispinus n.n.
scopa var. glomeratus n.n.: Note: Synonym of glomeratus.
scopa var. intermedius n.n.
scopa var. limiticola n.n.
scopa var. longispinus n.n.: Note: Synonym of rudibuenekeri.
scopa var. machadoensis Abraham: Succulenta 67 (4):81-84, 1988.
scopa var. maior n.n.
scopa var. major n.n.: Note: Synonym of floricomus var. spinosissimus ?
scopa var. marchesii Abraham: Kakt. u. a. Sukk. 40 (7):174-176, 1989.
scopa var. mayor n.n.: Note: Synonym of major.
scopa var. murielii n.n.: Prov. Desc.: A. Hofacker, Internoto 10 (2):40-42, 1989 and K. H. Prestlé, Internoto 14 (1):40-42, 1993
Note: Synonym of xicoi ?
scopa var. nivea n.n.
scopa var. nigrispinus n.n.
scopa var. prolifer n.n.
scopa var. ramosus (Osten) Backeberg: Notas sobre Cactáceas, obr. 18,Montevideo 1941
Die Cactaceae, 3:1637, Jena 1959.
scopa var. ramosus fa. albus n.n.: Note: Synonym of var. ruberrimus.
scopa var. ruberrimus n.n.
scopa var. rubrispinus n.n.
scopa var. splendens n.n.
scopa var. xicoi Abraham: Succulenta 67 (5):111-114, 1988.
scopa var. xiphacanthus Abraham: Succulenta 66 (12):256-260, 1987.
scopa fa. albilanatus Jahn: Stachelpost 4(22):75, 1969.
scopa fa. applanatus n.n.
scopa fa. candidus (Pfeiffer) Krainz: Enumeratio Diag. Cact., 64, Berlin 1837
Die Kakteen, C VIa, Stuttgart 1. 10. 1961.
scopa fa. daenikerianus (Krainz) Krainz: Schweizer Garten 7:199, 1945
Die Kakteen, C VIa, Stuttgart 1. 10. 1961.
scopa fa. erythrinus n.n.: Prov. Desc.: R. Havlicek, Internoto 9 (4):125-127, 1988.
scopa fa. erythrinus sbf. flavispinus n.n.
scopa fa. Fric
scopa fa. glauserianus (Krainz) Krainz [SP]: Schweizer Garten 7:201, 1945
Die Kakteen, C VIa, Stuttgart 1. 10. 1961.
scopa fa. Fechser: Prov. Desc.: H. Dopp, Internoto 10 (4):112-115, 1989.
And varieties from the following localities:: Erval, Rincao do Inferno and Sierra de los Canos.
scopa fa. Darilho: Note: Synonym of glomeratus.
And forms from the following localities:: Aigua, Bage, Cacapava, Casa de Pedra, Cerro Campanero, Erval, Garzon, Irapuazinho, Laguna Garzon, Lavalleja (Dept.), Maldonado, Marmaraja, Melo, Minas, MR 98, Pedras Altas, Pinheiro Machado, Piriapolis, Pororo, Procera, Punta Ballena, Santana de Boa Vista, Sierra Ballena and Rincon Segredo.
schlosseri Van Vliet: Succulenta 53 (1):10-13, 1974
Note: Transitional taxon between Scopanae and Mammulosi.
soldtianus Van Vliet: Succulenta 54 (4):72-75, 1975.
sucineus Ritter: Succulenta 49 (7):108-109, 1970.
sucineus var. albispinus n.n.
sucineus var. albus n.n.: Note: Synonym of var. albispinus ?
sucineus var. aureispinus n.n.: Note: Only a name.
sucineus var. caespitosus n.n.: Note: Only a name.
sucineus var. flavus n.n.: Note: Only a name.
sucineus var. flavispinus n.n.: Note: Only a name.
sucineus var. multicolorispinus n.n.: Note: Only a name.
sucineus var. sulphureus n.n.
And forms from the following localities:: Sao Gabriel.
sulphureus n.n.: Note: Synonym of sucineus var. sulphureus.

Transitional forms

With Mammulosi: fuscus, rauschii and schlosseri.

Section IV - Werdermannianae Havlicek

ferrugineus Schlosser: Internoto 3 (2):23-26, 1982.
memorialis Prestlé: Internoto 9 (1):3-7, 1988
Note: Transitional taxon between Werdermannianae and Herterianae via Herteri.
vanvlietii Rausch: Kakt. u. a. Sukk. 21 (5):89-90, 1970.
vanvlietii var. antonianus n.n.
vanvlietii var. gracilis Rausch: Kakt. u. a. Sukk. 21 (5):90, 1970.
vanvlietii var. gracilor ???
vanvlietii var. rubrispinus n.n.
And forms from the following localities:: Aqua Fria, Curtina, Tun. - Cla., Tun. - Quadr. and Tun. - Tacuarembo.
werdermannianus Herter: Rev. S. Am. Bot., 7:75-76, 1942
Note: Type species for Section IV.
werdermannianus var. albispinus n.n.
werdermannianus var. antonianus n.n.: Note: Synonym of vanvlietii var. antonianus.
werdermannianus var. brunispinus n.n.: Note: Provisional name for vanvlietii.
werdermannianus var. brunneispinus n.n.
werdermannianus var. caespitosus n.n.
werdermannianus var. concinnioides n.n.
werdermannianus var. grandiflorus n.n.
werdermannianus var. liliputanus n.n.
werdermannianus var. miniflorus n.n.
werdermannianus var. ruficeps n.n.: Note: Synonym of ferrugineus.
And forms from the following localities:: Cerro Porton, Quad., Quad. - Curtina and Paso del Cerro.

Transitional forms

With Herterianae: memorialis.

Section V - Setacei Fric

agnetae Van Vliet: Succulenta 54 (1):6-11, 1975.
agnetae var. aureispinus Van Vliet: Succulenta 54 (1):6-11, 1975.
agnetae var. minor Van Vliet: Succulenta 54 (1):6-11, 1975.
agnetae var. zapicanensis n.n.
amiguetii n.n.: Note: Provisional name for gibberulus.
apricus (Arechavaleta) Berger ex Backeberg: Flora Uruguaya, 5:205-207, Montevideo 1905
Kakteen, 211, Stuttgart 1929
Note: Type species for Section V.
apricus var. brevispinus n.n.: Note: Only a name.
apricus var. concinnus n.n.: Note: Synonym of concinnus.
apricus var. brunneispinus n.n.: Note: Synonym of concinnus var. brunneispinus.
apricus var. durispinus n.n.: Note: Synonym of concinnus var. durispinus.
apricus var. fimbriatiflorus n.n.: Note: Only a name.
apricus var. flaviflorus n.n.: Note: Only a name.
apricus var. grandiflorus n.n.
apricus var. multicostatus n.n.: Note: Synonym of multicostatus.
apricus var. muricatus n.n.: Note: Synonym of bommeljei.
apricus var. nigrispinus n.n.: Note: Synonym of concinnus var. nigrispinus.
apricus var. tabularis n.n.: Note: Synonym of tabularis.
And forms from the following localities:: Abra de Zavaleta, Cerro Arequita, MR 98, Pan de Azucar, Piriapolis, Pororo, Saulus, Serra das Minas and Sierra de Ballena.
arevaloensis n.n.
altasensis n.n.: Note: Provisional name gibberulus.
amiguetii n.n.: Note: Provisional name for gibberulus.
batoviensis n.n. [?]
blaauwianus Van Vliet: Succulenta 55 (6):108-113, 1976.
blaauwianus var. enormis Van Vliet: Succulenta 55 (6):108-113, 1976
Note: Synonym of multicostatus.
blaauwianus var. glaber ???
blaauwianus var. longispinus ???
blaauwianus var. nigrispinus ???
bommeljei Van Vliet: Succulenta 47 (1):5.8, 1968.
bommelianus n.n.: Note: Synonym of tabularis var. bommelianus.
brederooianus Prestlé: Succulenta 64 (4):84-87, 1985.
concinnioides n.n.: Prov. Desc.: W. Prauser, Internoto 14 (2):53-58, 1993.
concinnus (Monville) Berger ex Backeberg: l'horticulteur universel, 222, 1839
Kakteen, 210, Stuttgart 1929.
concinnus var. apricus n.n.: Note: Synonym of apricus.
concinnus var. aceguaensis Gerloff: Internoto 10 (4):99-106, 1989.
concinnus var. arevaloensis n.n.: Note: Synonym of arevaloensis.
concinnus var. ballenaensis n.n.
concinnus var. brunneispinus n.n.
concinnus var. cunapiruensis n.n.: Note: Provisional name for rubrigemmatus.
concinnus var. durispinus n.n.: Prov. Desc.: Wolfgang Prauser, Internoto 17 (2):44-46, 1996.
concinnus var. elegans n.n.
concinnus var. eremiticus (Ritter) Gerloff: Kakt. in Süd., 1:180-181, Spangenberg 1979
Internoto 9 (4):99-112, 1988.
concinnus var. fimbriatiflorus n.n.
concinnus var. joadii (Hooker) Ito: Curtis's Botanical Magazine, 112, 1886
Explan. Diagr. Austroechinocactinae, 243, 1957.
concinnus var. gerbalitoensis n.n.: Note: Misspelling of concinnus var. yerbalitoensis.
concinnus var. gibberulus (Prestlé) Gerloff: Succulenta 65 (6-7):142-145, 1986
Internoto 9 (4):99-112, 1988.
concinnus var. longispinus n.n.
concinnus var. luysianus n.n. [ST]: Note: Misspelling of wolthuysianus.
concinnus var. montevidensis n.n.
concinnus var. mosquitensis n.n.: Note: Synonym of megalanthus.
concinnus var. multicostatus n.n.: Note: Synonym of multicostatus.
concinnus var. nigrispinus n.n.
concinnus var. olimarensis n.n.: Note: Synonym of olimarensis.
concinnus var. parviflorus n.n.
concinnus var. piriapolisensis n.n.
concinnus var. polancoensis n.n.
concinnus var. pororensis n.n.: Prov. Desc.: Marc Dumon, Internoto 17 (3):100-101, 1996.
concinnus var. quervosensis n.n.
concinnus var. rubrigemmatus (Abraham) Gerloff: Kakt. u. a. Sukk. 39 (2):38-41, 1988
Internoto 9 (4):99-112, 1988.
concinnus var. wolthuysianus n.n. [ST]: Note: Provisional name for blaauwianus.
concinnus var. yerbalitoensis n.n.: Prov. Desc.: Internoto 12 (2):58-62, 1991.
concinnus var. yubaldensis n.n.
And varieties/forms from the following localities:: Animas, Arevelo, Casupa, Carlos, Cerro de Montevideo, Parador, Picada Benitez and Trinidad.
concinnus fa. Cunapriú fa. Cunapriú: Note: Synonym of rubrigemmatus.
concinnus fa. MR98 "nigrus" ???
elegans n.n.: Note: Synonym of tabularis var. elegans or concinnus var. elegans.
eremiticus: Note: Reduced to a variety of concinnus
Note: arnostianus has also been distributed under this name.
gibberulus: Note: Reduced to a variety of concinnus.
islaensis n.n.
joadii: Note: Synonym of concinnus var. joadii.
mannii: Note: Synonym of tabularis.
multicostatus Buining et Brederoo: Die Kakteen, C VIc, Stuttgart 3. 12. 1972.
multicostatus var. umbrinispinus n.n.
multicostatus var. prolifer n.n.: Note: Only a name.
neoblaauwianus n.n. [?]
nigroleptacanthus n.n.
notabilis n.n.
obscurus n.n.: Prov. Desc.: Karl Heinz Prestlé, Internoto 17 (2):66-69, 1996
olimarensis n.n.
parviflorus n.n.: Note: Synonym of concinnus var. parviflorus.
piriapolisensis n.n.: Note: Synonym of concinnus var. piriapolisensis
polancoensis n.n.: Note: Synonym of concinnus var. polancoensis
pseudoblaauwianus n.n. [?]
pseudoblaauwianus var. nigrispinus n.n. [?]
rubricurvispinus n.n.
rubrigemmatus: Note: Reduced to a variety of concinnus.
seticeps n.n.: Note: Provisional name for agnetae.
seticeps var. minor n.n.: Note: Provisional name for agnetae var. minor.
spinibarbis Ritter [?]: Kakt. in Süd., 1:186-187, Spangenberg 1979.
tabularis (Cels ex Rümpler) Berger ex Backeberg: Handbuch der Cacteenkunde, 552, Leipzig 1886
Kakteen, 211, Stuttgart 1929.
tabularis var. bommelianus n.n.
tabularis var. elegans n.n.
tabularis var. major n.n.
tabularis var. nigrispinus n.n.
tabularis var. setispinus n.n.
tabularis var. splendens n.n.
tabularis var. velenovskyi n.n.
And forms from the following localities:: Piriapolis, Punta Ballena, San Antonio, Sierra de las Animas and Tolomban.
setispinus n.n.: Note: Synonym of tabularis var. setispinus.
veerbekianus n.n.
villaseranus n.n. [?]

Transitional forms

With Mammulosi: megalanthus.

Section VI - Uebelmannianae Havlicek

arachnites Ritter: Succulenta 49 (7):108-109, 1970.
arachnites var. gracilior n.n.: Note: Synonym of crassigibbus var. gracilior.
arachnites var. minor Ritter: Succulenta 49 (7):108-109, 1970.
arachnites var. minor fa. darilhoensis Gerloff: Internoto 18 (4):119-127, 1997.
arachnites var. parviflorus n.n.
arachnites fa. Irapuazinho
crassigibbus Ritter: Succulenta 49 (7):108-109, 1970.
crassigibbus var. difformis n.n.: Prov. Desc.: R. Havlicek, Internoto 3 (2):30, 1982.
crassigibbus var. gracilior n.n.
crassigibbus var. grandiflorus n.n.
crassigibbus var. miniflorus n.n.: Note: Only a name ?
crassigibbus var. minor n.n.
crassigibbus var. polyacanthus n.n.: Prov. Desc.: R. Havlicek, Internoto 3 (2):30, 1982.
And forms from the following localities:: Itaoca and Pinheiro Machado.
darilhoensis n.n.: Note: Synonym of arachnites var. minor fa. darilhoensis
meonacanthus Prestlé: Internoto 7 (2):35-39, 1986.
parviflorus n.n.: Note: Synonym of uebelmannianus fa. flaviflorus or arachnites var. parviflorus ?
pleiocephalus n.n.: Note: Synonym of uebelmannianus var. pleiocephalus.
uebelmannianus Buining: Kakt. u. a. Sukk. 19 (9):175-176, 1968
Note: Type species for Section VI.
uebelmannianus var. nilsonii Königs: Internoto 11 (4):119-124, 1990.
uebelmannianus var. pleiocephalus Gerloff et Königs: Internoto 13 (4):103-112, 1992.
uebelmannianus var. pleiocephalus fa. flaviflorus: Note: This is a Homonym of uebelmannianus fa. flaviflorus so was renamed fa. gilviflorus.
uebelmannianus var. pleiocephalus fa. gilviflorus Gerloff et Königs: Internoto 13 (4):103-112, 1992
Internoto 14 (2):70, 1993.
And varieties from the following localities:: Santana da Boa Vista.
uebelmannianus fa. flaviflorus Buining: Kakt. u. a. Sukk. 19 (9):175-176, 1968.
uebelmannianus fa. Encruzilhade: Note: Synonym of uebelmannianus var. nilsonii.
uebelmannianus fa. "Sprossend": Note: Synonym of uebelmannianus var. pleiocephalus.
And forms from the following localities:: Estancia Tala.
werneri (Buining) Hofacker: Note: The IOS consensus name for uebelmannianus.

Section VII - Herterianae Havlicek

callianthus n.n.: Prov. Desc.: R. Havlicek, Internoto 4 (2):49, 1983.
cupreatus n.n.: Note: Provisional name for roseoluteus.
erythrinus n.n.: Note: Synonym of horstii var. purpureiflorus.
herteri (Werdermann) Werdermann, Buining et Kreuzinger: Rev. S. Am. Bot., 3 (4/6):143-144, 1936
Succulenta 29 (2):17-19, 1950
Note: Type species for Section VII.
herteri var. albispinus n.n.
herteri var. aurisetus n.n.
herteri var. erytranthus n.n.: Note: Misspelling of erythrinus ?
herteri var. "fechseri" ???
herteri var. luteo-roseus n.n.: Note: Synonym of roseoluteus ?
herteri var. porphyreus n.n.
herteri var. pseudoherteri (Buining) Herm: National Cactus and Succulent Journal 26 (1):2-3, 1971
Internoto 10 (2):35-39, 1989.
And forms from the following localities:: Rincao da Bolsa.
horstii Ritter: Succulenta 45 (1):3-4, 1966.
horstii var. erythrinus n.n.: Note: Synonym of horstii var. purpureiflorus.
horstii var. mugelianus n.n.: Note: Synonym of mugelianus.
horstii var. purpureiflorus Ritter: Kakt. in Süd., 1:184, Spangenberg 1979.
horstii fa. mugelianus Herm: Internoto 14 (2):59-64, 1993.
horstii fa. purpureus (Ritter) Herm: Succulenta 49 (7):108-109, 1970
Internoto 14 (2):59-64, 1993.
muegelianus n.n.: Note: Misspelling of mugelianus.
mugelianus n.n.: Note: Reduced to a form of horstii.
pseudoherteri: Note: Reduced to a variety of herteri.
purpureus: Note: Reduced to a form of horstii.
purpureus var. mugelianus n.n.: Note: Synonym of mugelianus.
purpureus fa. Waras
roseoluteus Van Vliet: Succulenta 52 (6):108-113, 1973
Note: Transitional taxon between Herterianae and Mammulosi.
roseoluteus var. flaviflorus n.n. ???
roseoluteus var. roseiflorus n.n.
rubriflorus Kolischer: Verzeichnis amerikanischer und anderer Sukkulenten mit Revision der Systemik der Kakteen, Eger 1935
Note: Synonym of herteri.

Transitional forms of Notocactus and Neonotocactus

With Mammulosi: arnostianus, curvispinus and ritterianus.

Subgenus Neonotocactus Buxbaum

Section VIII - Mammulosi Fric

alamoensis n.n.
allosiphon Marchesi: Bol. Soc. Arg. Bot. 14 (3):246-248, 1972
Note: Transitional taxon between Mammulosi and Malacocarpus.
allosiphon var. albiareolatus n.n.
allosiphon var. albispinus n.n. ???
allosiphon var. pulchellus n.n.
allosyphon: Note: Misspelling of allosiphon.
arnostianus Kolarik et Lisal: Internoto 7 (1):6-19, 1986
Note: Transitional taxon between Neonotocactus and Notocactus.
brasiliensis n.n.: Note: Synonym of mammulosus var. brasiliensis.
brasiliensis var. rubrispinus n.n.: Note: Synonym of mammulosus var. rubrispinus.
buiningii Buxbaum: Kakt. u. a. Sukk. 19 (12):229-231, 1968
Note: Transitional taxon between Mammulosi and Malacocarpus.
buiningii var. alamoensis n.n.: Note: Synonym of alamoensis.
buiningii var. brunneispinus n.n.
buiningii var. ferrugineispinus n.n.: Note: Synonym of ferrugineus.
buiningii var. idiotensis n.n.: Note: Synonym of mammulosus var. gracilispinus ?
buiningii var. pulchellus n.n.: Note: Synonym of allosiphon var. pulchellus.
buiningii var. schlosseri n.n.
And forms from the following localities:: Catalan Grande.
catalanensis n.n.
cristatoides Ritter: Kakt. in Süd., 1:190-191, Spangenberg 1979.
curvispinus Ritter: Kakt. in Süd., 1:189, Spangenberg 1979
Note: Transitional taxon between Neonotocactus and Notocactus
Note: Both arnostianus and ritterianus have also been distributed under this name.
elegans n.n.: Note: A name used for arnostianus, curvispinus and ritterianus.
erubescens (Osten) Marchesi: Notas sobre Cactáceas, 47-50, Montevideo 1941
Bol. Soc. Arg. Bot., 14 (3):246-248, 1972.
erythracanthus Schlosser et Brederoo: Kakt. u. a. Sukk. 36 (9):186-189, 1985.
euvelenovskyi Fleischer et Schütz: Friciana 8 (50):24-25,36, 1975.
fasciculatus n.n.: Note: Synonym of orthacanthus.
floricomus (Arechavaleta) Berger ex Backeberg: Flora Uruguaya, 5:183-185, Montevideo 1905
Kakteen, 211, Stuttgart 1929.
floricomus var. flavispinus Backeberg: Cactus and Succulent Journal of America 23:85, 1951.
floricomus var. grandiflorus n.n.: Note: Synonym of erythracanthus Schl. 165.
floricomus var. multiflorus n.n.: Note: Synonym of erythracanthus Schl. 165a.
floricomus var. rubrispinus n.n.: Note: Synonym of erythracanthus Schl. 165b.
floricomus var. rutilans n.n. [ML]: Note: Provisional name for rutilans.
floricomus var. spinosissimus Fric ex Buining: Succulenta 36 (7):77-84, 1957.
floricomus var. spinosissimus fa. robusta n.n.: Note: Only a name ?
floricomus var. velenovskyi Krainz: Schweizer Garten 6:225, 1947.
And forms from the following localities:: Rivera and Tacuarembo.
macambarensis Prestlé: Internoto 7 (3):67-72, 1986.
mammulosus (Lemaire) Berger ex Backeberg: Cact. aliq. nov. ins. Hort. Monv., 40, Paris 1838
Kakteen, 212, Stuttgart 1929
Note: Type species for the Subgenus and Section VIII.
mammulosus var. albispinus n.n.
mammulosus var. albiarmatus n.n.
mammulosus var. "andreae" n.n. ???
mammulosus var. arapeyensis n.n.
mammulosus var. arbolitoensis n.n.
mammulosus var. arevaloensis n.n.
mammulosus var. artinensis n.n.
mammulosus var. brasiliensis Havlicek: Kaktusy, 16(1):5-7, 1980.
mammulosus var. brasiliensis fa. meldiansis n.n.
mammulosus var. buiningioides n.n.
mammulosus var. catalanensis n.n.: Note: Synonym of catalanensis.
mammulosus var. elegans n.n.
mammulosus var. erubescens n.n.: Note: Synonym of erubescens.
mammulosus var. floricomus n.n.
mammulosus var. floricomus fa. erubescens n.n.
mammulosus var. gracilior n.n.
mammulosus var. gracilis n.n.: Note: Misspelling or synonym of var. gracilispinus.
mammulosus var. gracilispinus n.n.
mammulosus var. grandiflorus n.n.
mammulosus var. hircinus n.n.: Note: Only a name.
mammulosus var. intricatissimus n.n.
mammulosus var. longispinus Fric
mammulosus var. luteospinus n.n.
mammulosus var. machadoensis n.n.
mammulosus var. macrocanthus n.n.: Note: Synonym of mammulosus var./fa. PR 88.
mammulosus var. marmarajensis n.n.
mammulosus var. masollerensis n.n.
mammulosus var. minor n.n.: Note: Only a name.
mammulosus var. multiflorus n.n.
mammulosus var. nigrispinus n.n.
mammulosus var. palomaensis n.n.: Note: Synonym of erythracanthus PR 82.
mammulosus var. paucicostatus n.n.
mammulosus var. piriapolisensis n.n.
mammulosus var. rubrispinus n.n.
mammulosus var. rubrispinus fa. robustior n.n.
mammulosus var. sarandiensis n.n.: Note: Also seen as sarandinensis but if this taxon comes from Sarandi then sarandiensis is the correct name.
mammulosus var. spinosior n.n.: Note: Only a name.
mammulosus var. tunuayensis n.n.
mammulosus var. uruguayensis n.n.
mammulosus var. Ancasti: Note: Synonym of submammilatus (Lau 502).
mammulosus fa. Catalan Grande: Note: Synonym of catalanensis.
mammulosus fa. / var. uruguayanus n.n.
And forms from the following localities:: Alamo, Artigas, Bage, Balve, Bossoroca, Canada de los Buros, Bavera, Colonia Itapeti, Colonia Pintado, Cuchilla Areicua, Cuchilla Negra, Espinilho, Hiberalta, Lavras do Sul, Livramento, Minas de Corrales, Paso Ricardinho, Quarai, Rincao do Inferno, Rivera, Tranqueras, Treinta-y-Tres, Uruguay, Uruguay-Colonia, Vale do Platao, Varzea and Vichadero.
megalanthus Schlosser et Brederoo: Kakt. u. a. Sukk. 32 (7):154-157, 1981
Note: Transitional taxon between Mammulosi and Setacei.
meldiansis n.n.: Note: Synonym of brasiliensis fa. meldiansis.
notabilis n.n.
orthacanthus (Link et Otto) Van Vliet: Verh. Ver. Beförd. Gartenb., 3:427, Berlin 1827
Succulenta 49 (11):185-190, 1970
Note: Transitional taxon between Mammulosi and Malacocarpus.
orthacanthus var. minor ???
pampeanus Spegazzini: Contribucion al estudio de la flora de la Sierra de la Ventana, 27-28, La Plata 1896
Note: Synonym of submammulosus var. pampeanus.
permutatus Ritter: Kakt. in Süd., 1:188-189, Spangenberg 1979.
pseudoeremiticus n.n.: Note: Synonym of arnostianus.
ritterianus Kolarik et Lisal: Internoto 7 (1):3-5 9-19, 1986
Note: Transitional taxon between Neonotocactus and Notocactus.
rubropedatus Ritter: Kakt. in Süd., 1:189-190, Spangenberg 1979.
sp. Entre Rios: Note: Synonym of turecekianus.
spinosissimus n.n.: Note: Synonym of floricomus var. spinosissimus.
submammilatus n.n.: Note: Synonym of submammulosus ssp. minor.
submammulosus (Lemaire) Backeberg: Cactearum genera nova speciesque novae et omnium in Horto Monvilliano cultarum, 20, Paris 1839
Kaktus ABC, 255, Kopenhagen 1935.
submammulosus var. elegans n.n. ???
submammulosus ssp. minor (Kiesling) Abraham: Cactus and Succulent Journal (US), 67:14-22, 1995
Internoto 16 (3):82-86, 1995.
submammulosus var. longispinus n.n.
submammulosus var. pampeanus (Spegazzini) Backeberg: Contribucion al estudio de la flora de la Sierra de la Ventana, 27-28, La Plata 1896
Die Cactaceae, 3:1649-1650, Jena 1959.
submammulosus fa. tunuayensis n.n.
And forms from the following localities:: Dept. San Jose, Charrua, Cordoba, Los Funes, Paso del Toros and Tacuarembo.
turecekianus (Kiesling) Abraham: Cactus and Succulent Journal (US), 67:14-22, 1995
Internoto 16 (3):82-86, 1995.
velenovskyi n.n.: Note: Synonym of euvelenovskyi.

Transitional forms

With Scopanae: fuscus, rauschii and schlosseri.
With Herterianae: roseoluteus.
With Malacocarpus: allosiphon, buiningii, nivosus and orthacanthus.
With Notocactus: arnostianus, curvispinus and ritterianus.
With Melchersianae: mammulosus var. curtinensis.

Section IX - Melchersianae Havlicek

curtinensis n.n.: Note: Synonym of mammulosus var. curtinensis.
engleri n.n.: Note: Synonym or Misspelling of eugeniae.
erizo n.n.: Note: Synonym of mueller-moelleri ?
eugeniae Van Vliet: Succulenta 55 (2):24-26, 1976.
eugeniae var. microfloridus n.n.: Note: Local form of eugeniae with a close relationship to veenianus.
eugeniae var. minimus n.n.: Note: Provisional name for veenianus.
gutierrezii Abraham: Kakt. u. a. Sukk. 39 (7):150-154, 1988.
mammulosus var. curtinensis n.n.: Note: Transitional taxon between Mammulosi and Melchersianae
Prov. Desc.: R. Havlicek, Internoto 5 (1):12-13, 1984.
mueller-melchersii Fric ex Backeberg: Kaktus ABC, 254, 415, Kopenhagen 1935
Note: Type species for Section IX.
mueller-melchersii var. colorispinus n.n.: Prov. Desc.: R. Havlicek, Internoto 5 (1):10, 1984.
mueller-melchersii var. elongatus n.n.: Prov. Desc.: S. Theunissen, Internoto 11 (2/3):104, 1990.
mueller-melchersii var. elegans n.n.: Note: Provisional name for winkleri.
mueller-melchersii var. ferrugineus n.n.: Prov. Desc.: R. Havlicek, Internoto 5 (1):10, 1984.
mueller-melchersii var. longispinus n.n.: Note: A name also used by the German firm SPI, Wiesbaden for erizo.
mueller-melchersii var. longispinus fa. Ruta 8/km 357
mueller-melchersii var. multicolorispinus n.n.: Note: Synonym of var. colorispinus.
mueller-melchersii var. tacuarembensis n.n.: Prov. Desc.: R. Havlicek, Internoto 5 (1):11, 1984.
mueller-melchersii var. nigrispinus n.n.
mueller-melchersii var. rosea n.n.: Note: Synonym of winkleri.
mueller-melchersii var. roseus n.n.: Note: Synonym of var. ferrugineus.
mueller-melchersii fa. gracilispinus (Krainz) Krainz: Kakteenkunde 2:33-35, 1942
Die Kakteen, C VIa, Stuttgart 1. 1. 1962.
mueller-melchersii fa. gracilispinus sbf. nigrispinus n.n.: Note: Synonym of var. nigrispinus.
mueller-moelleri Fric ex Fleischer et Schütz: Friciana 8 (50):26-30, 36, 1975.
mueller-moelleri var. albispinus n.n. ???
mueller-moelleri var. caveraensis n.n.
mueller-moelleri var. gracilispinus n.n. ???
And forms from the following localities:: Bage and Varzea.
paulus Schlosser et Brederoo: Kakt. u. a. Sukk. 31 (4):114-116, 1980.
paulus var. longispinus ???
pseudorutilans n.n.: Note: Provisional name for eugeniae.
pseudorutilans var. microfloridus n.n.: Note: Provisional name for veenianus.
pseudorutilans var. minimus n.n.: Note: Provisional name for veenianus.
roseiflorus Schlosser et Brederoo: Kakt. u. a. Sukk. 29 (12):273-277, 1978.
rubrispinus n.n.
rutilans Däniker et Krainz: Sukkulentenkunde 2:19, 1948.
rutilans var. jasminiflorus n.n.: Note: Provisional name for roseiflorus.
rutilans var. grandiflorus n.n.: Note: Only a name ?
rutilans var. longispinus n.n.
rutilans var. microfloridus n.n.
rutilans var. rhodacanthus n.n.: Note: Provisional name for veenianus.
rutilans var. rubrispinus n.n.: Note: Synonym of erythracanthus.
rutilans fa. storianus Pazout: Kaktusy 66 2 (6):125-126, 1966.
sp. Rio Cuaré [ML]: Note: Misspelling of Rio CuaróCuaró.
sp. Rio Cuaró . [ML]: Note: Synonym of paulus.
veenianus Van Vliet: Succulenta 53 (9):171-173, 1974.
winkleri Van Vliet: Succulenta 54 (7):136-139, 1975.
ycarensis n.n.: Note: Synonym of roseiflorus.
zonalis n.n.: Note: Synonym of mueller-melchersii var. colorispinus.

Transitional forms

With Mammulosi: mammulosus var. curtinensis.

Subgenus Malacocarpus (Salm Dyck) Buxbaum

aciculatus n.n.: Note: Provisional name for orthacanthus.
acuatus (Link et Otto) Theunissen: Verh. Ver. Beförd. Gartenb., 3:424, Berlin 1827
Succulenta 60 (6):140-142, 1981.
acuatus var. depressus: Note: Transferred to a variety of tephracanthus.
acuatus var. prolifer Ritter: Note: Synonym of prolifer.
albiflorus n.n.
attenuata n.n.
beltranii (Fric ex Fleischer et Schütz) Schäfer [MC]: Friciana 8 (50):33-34, 36-37, 1975
Kakteen-Sukkulenten 14 (1-4):36, 1979.
bertramii: Note: Misspelling of beltranii ?
bezrucii (Fric) Schäfer: Praktický rádce 17 (7,8):100, 110, 1922
Kakteen-Sukkulenten 14 (1-4):36-37, 1979.
bezrucii var. centrispinus (Fric) Theunissen: Praktický rádce 17 (7):110, 1922
Succulenta 60 (6):140-142, 1981.
bezrucii var. corniger (Fric) Theunissen: Praktický rádce 17 (7):110, 1922
Succulenta 60 (6):140-142, 1981.
calvescens Gerloff et Nilson: Internoto 15 (3):75-83, 1994.
cephalophorus n.n.
congregatus n.n.: Prov. Desc.: K. H. Prestlé, Internoto 5 (4):138-140, 1984.
corynodes: Note: Reduced to a form of erinaceus.
courantii: Note: Reduced to a variety of sellowii.
curtinensis n.n.
dalvae n.n.
discophalium n.n.: Note: Provisional name of neohorstii var. prestlei.
erinaceus (Haworth) Krainz: Supplementum Plantarum Succulentarum, 74, London 1819
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
erinaceus fa. corynodes (Otto ex Pfeiffer) Gerloff: Enumeratio Diag. Cact., 55, Berlin 1837
Notokakteen, 152-153, 1995.
Note: Type species for the Subgenus
erinaceus var. kovarikii n.n.: Note: Synonym of kovarikii.
erinaceus var. leucocarpus n.n.: Note: Synonym of leucocarpus.
erinaceus var. macrocanthus n.n.: Note: Synonym of macrocanthus.
erinaceus var. spinosior n.n.: Note: Synonym of tephracanthus.
ernestii n.n: Note: Synonym of pauciareolatus.
flaviflorus n.n.: Note: Synonym of albiflorus.
fricii: Note: Reduced to a form of sellowii.
fricii var. robustior n.n.: Note: Only a name ?
gladiatus n.n.
glochidiatus n.n.: Note: Misspelling of gladiatus ?
grandicornis n.n.
hamesterii n.n.
hennisii n.n.: Note: Only a name.
horridispinus n.n.
infernensis n.n.: Note: Synonym of dalvae ?
Prov. Desc.: K. H. Prestlé, Internoto 5 (3):98-99, 1984.
infernensis fa. atrispinus n.n.: Prov. Desc.: K. H. Prestlé, Internoto 5 (3):99, 1984.
intricatus (Link et Otto) Herter: Verh. Ver. Beförd. Gartenb., 3:428, Berlin 1827
Cactus 120, 1954.
invictus n.n.
kovarikii (Fric ex Berger) Krainz: Kakteen, 206, Stuttgart 1929
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
langsdorfii (Lehmann) Krainz: Index seminum hort. Hamburg, 17, 1827
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
langsdorfii ssp. multiceps Hofacker et Herm: KuaS, 50(3):62-64, 1999.
langsdorfii ssp. pulvinatus (Van Vliet) Hofacker et Herm: Succulenta 49 (4):50-52, 1970
KuaS 50(3):62, 1999
langsdorfii var. leprosorum (Ritter) Hofacker: Kakt. in Süd., 1:194-195, Spangenberg 1979
Internoto 14 (4):124, 1993.
langsdorfii var. longispinus (Ritter) Gerloff: Kakt. in Süd., 1:198-199, Spangenberg 1979
Internoto 14 (2):78-79, 1993.
langsdorfii fa. eddiewarasii n.n.: Prov. Desc.: N. Gerloff and G. Königs, Internoto 14 (2):78, 1993.
langsdorfii fa. prolifer (Ritter) Gerloff: Kakt. in Süd., 1:199, Spangenberg 1979
Internoto 14 (2):76-77, 1993.
leprosorum: Note: Reduced to a variety of langsdorfii.
leucocarpus: Note: Reduced to a form of sellowii.
longispinus: Note: Reduced to a variety of langsdorfii.
macrocanthus: Note: Reduced to a form of sellowii.
macrogonus: Note: Reduced to a form of sellowii.
maldonadensis: Note: Reduced to a variety of neoarechavaletae.
neoarechavaletae (Schumann ex Spegazzini) Havlicek: Cact. Plat. Tentamen, 494, Buenos Aires 1905
Kakt. Vilag 18 (4):79, 1988.
neoarechavaletae var. kovarikii n.n.: Note: Synonym of kovarikii.
neoarechavaletae var. marchesii n.n.
neoarechavaletae var. maldonadensis (Herter ex Herter) Havlicek: Florula Uruguayensis,90, Montevideo 1930
Internoto 14 (4):126-127, 1994.
neoarechavaletae fa. nigrispinus n.n.
And forms from the following localities:: Garzon, Punta ballena, Treinta y Tres and Tupambae,.
neohorstii (Ritter) Theunissen: Kakt. in Süd., 1:199-200, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
neohorstii var. juvenaliformis (Ritter) Theunissen: Kakt. in Süd., 1:200, Spangenberg 1979
Succulenta 60 (6):140-142, 1981
Note: Reduced to a form.
neohorstii var. prestlei Gerloff: Internoto 16 (1):3-17, 1995.
neohorstii fa. juvenaliformis (Ritter) Gerloff: Kakt. in Süd., 1:200, Spangenberg 1979
Internoto 14 (2):73, 1993.
neohorstii fa. ravenditii n.n.
nivosus n.n.: Note: Transitional taxon between Malacocarpus and Mammulosi
Prov. Desc.: K. H. Prestlé, Internoto 5 (2):59-61, 1984.
nivosus var. albiflorus n.n.: Prov. Desc.: K. H. Prestlé, Internoto 13 (2):46-49, 1992.
nivosus var. clavarioides n.n.
nivosus var. gracilispinus n.n.
oreophilus n.n.
"patagonicus": Note: Synonym of Austrocactus patagonicus
pauciareolatus: Note: Reduced to a form of sellowii.
paucicostatus n.n.
polyacanthus (Link et Otto) Theunissen: Verh. Ver. Beförd. Gartenb., 3:422, Berlin 1827
Succulenta 60 (6):140-142, 1981
Note: Synonym of langsdorfii.
prolifer: Note: Reduced to a form of langsdorfii.
pseudopulvinatus n.n.: Note: Synonym of pulvinatus ?
pulvinatus: Note: Reduced to a subspecies of langsdorfii.
rubricostatus: Note: Reduced to a form of sellowii.
schaeferianus (Abraham et Theunissen) Havlicek: Internoto 9 (1):21-25, 1988
Internoto 9 (2):39, 1988
Note: Synonym of turbinatus ?
sellowii (Link et Otto) Theunissen: Verh. Ver. Beförd. Gartenb., 3:425, Berlin 1827
Succulenta 60 (6):140-142, 1981.
sellowii var. courantii (Lemaire) Gerloff: Cact. aliq. nov. ins. Hort. Monv., 20, Paris 1838
Internoto 14 (2):73, 1993.
sellowii var. turbinatus (Arechavaleta) Havlicek: Flora Uruguaya, 5:235-237, Montevideo 1905
Internoto 15 (3):89, 1994.
sellowii fa. fricii (Arechavaleta) Gerloff: Flora Uruguaya, 5:244-246, Montevideo 1905
Notokakteen, 163, 1995.
sellowii fa. leucocarpus (Arechavaleta) Gerloff: Flora Uruguaya, 5:239-242, Montevideo 1905
Notokakteen, 164, 1995.
sellowii fa. macrocanthus (Arechavaleta) Gerloff: Flora Uruguaya, 5:230-232, Montevideo 1905
Internoto 14 (2):72, 1993.
sellowii fa. macrogonus (Arechavaleta) Havlicek: Flora Uruguaya, 5:232-234, Montevideo 1905
Internoto 15 (3):89, 1994.
sellowii fa. pauciareolatus (Arechavaleta) Gerloff: Flora Uruguaya, 5:246-247, Montevideo 1905
Internoto 14 (2):72-73, 1993.
And forms from the following localities:: Catalan Grande, Cerro Accolorado, Cerro de Montevideo, Cuchilla Areicua, Cunapiru, Laguna Garzon and Punta de la Arena.
sellowii fa. rubricostatus (Fric ex Fleischer et Schütz) Gerloff: Friciana 8 (50):37, 1975
Notokakteen, 165, 1995.
sessiliflorus (Hooker) Krainz: Curtis's Botanical Magazine, 64, 1837
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
Note: Backeberg (Die Cactaceae, 3:1621, Jena 1959) and Britton and Rose (The Cactaceae 3:188, 1920) both state that Mackie was the original author.
sessiliflorus var. debilacanthus n.n.
sessiliflorus var. martinii (Labouret ex Rümpler) Krainz: Handbuch der Cacteenkunde, 454-455, Leipzig 1886
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
sessiliflorus var. pauciareolatus n.n.
sessiliflorus var. stegmannii (Backeberg) Havlicek: Die Cactaceae, 3:1623, Jena 1959
Internoto 15 (4):129, 1994.
And forms from the following localities:: Saavedra, Sierra de la Ventana and Sierra de Tandil.
stegmannii: Note: Reduced to a variety of sessiliflorus.
tephracanthus (Link et Otto) Krainz: Verh. Ver. Beförd. Gartenb., 3:422, Berlin 1827
Kakt. u. a. Sukk. 17 (10):195-196, 1966.
tephracanthus var. depressus (Spegazzini) Stuchlík: Cact. Plat. Tentamen, 494, Buenos Aires 1905
Rod Notocactus 147, 1993.
tetracanthus n.n.
theunissianus n.n.
tricornis n.n.
turbinatus: Note: Reduced to a variety of sellowii.
viridiflavus n.n.
vorwerkianus (Werdermann) Krainz: Neue Kakteen, 101-102, Frankfurt 1931
Kakt. u. a. Sukk. 17 (10):195-196, 1966
Note: A variety of sessiliflorus ?

Transitional forms

With Mammulosi: allosiphon, buiningii, nivosus and orthacanthus.

Subgenus Brasilicactus (Backeberg) Buxbaum

Note: Acanthocephala Backeberg was also a name used for this subgenus.

elachisanthus (Weber) Herter: Bull. Mus. Hist. Nationale de Paris X, 387, 1904
Cactus 120, 1954.
graessneri (Schumann) Berger ex Krainz: Monatsschrift für Kakteenkunde, 13:130, 1903
Kakteen, 208-209, Stuttgart 1929.
graessneri var. albisetus (Cullmann) Krainz: Kakt. u. a. Sukk. 6 (6):105, 1955
Die Kakteen, C VIa, Stuttgart 15. 7. 1960.
graessneri var. aureiflorus n.n.
graessneri var. flaviflorus Backeberg [BC]: Das kakteenlexicon 76, 1977.
graessneri var. fortalezensis n.n.
graessneri var. jaquiranensis n.n.
graessneri var. longispinus n.n.
graessneri var. luteus n.n.
graessneri var. stellatus Hort. ex Backeberg Reference ???
graessneri var. tenuispinus n.n.
graessneri var. vacariensis n.n.: Note: Synonym of var. stellatus.
graessneri fa. fortalezensis Prestlé Reference ???
graessneri fa. microdasys Braun: Kakt. u. a. Sukk. 37 (3):57, 1986.
And forms from the following localities:: Morro de Chapeu.
haselbergii (Haage) Berger ex Krainz: Handbuch der Cacteenkunde, 563-564, Leipzig 1886
Kakteen, 208, Stuttgart 1929
Note: Type species for the Subgenus.
haselbergii var. albatus n.n.
haselbergii var. albolanatus n.n.
haselbergii var. gracilispinus n.n.
haselbergii var. lanatus n.n.
haselbergii var. luteus n.n.
haselbergii var. pseudograessneri n.n.
haselbergii var. robustispinus n.n.: Note: Synonym of graessneri var. stellatus ?
haselbergii var. splendens n.n.: Note: Only a name ?
haselbergii var. stellatus n.n.: Note: Synonym of graessneri var. stellatus.
haselbergii var. vacariensis n.n.: Note: Synonym of graessneri var. stellatus.

Subgenus Eriocactus (Backeberg) Buining

Note: Eriocephalus Backeberg and Eriocephala (Backeberg) Backeberg were also names used for this subgenus.

ampliocostatus (Ritter) Theunissen: Kakt. in Süd., 1:253-255, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
claviceps (Ritter) Krainz: Succulenta 45 (8):115-116, 1966
Katalog der Städtischen Sukkulenten - Sammlung, 89, Zürich 1967.
grossei (Schumann) Fric: Monatsschrift für Kakteenkunde, 9:44, 1899
Cacti the coming fashion, Praha 1928.
grossei var. aureispinus (Ritter) Theunissen: Kakt. in Süd., 1:253, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
itapeitiensis Mori et Ito: Note: Only a name ?
leninghausii (Haage Sen.) Berger ex Krainz: Monatsschrift für Kakteenkunde, 5:147, 1895
Kakteen, 209, Stuttgart 1929.
leninghausii var. longispinus n.n.: Note: Synonym of var. minor.
leninghausii var. minor (Ritter) Theunissen: Kakt. in Süd., 1:156, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
leninghausii var. nigrispinus n.n.: Note: Only a name.
leninghausii fa. apelii (Heinrich) Krainz: Kakteenkunde, 1-3, 1940
Die Kakteen, C VIc, Stuttgart 1. 7. 1968.
magnificus (Ritter) Krainz: Kakt. u. a. Sukk. 17 (10):195-196, 1966
Succulenta 45 (4):50-52, 1966.
nigrispinus (Schumann) Buining: Monatsschrift für Kakteenkunde, 9:44, 1899
Succulenta 49 (11):172-181, 1970.
schumannianus (Nicolai) Fric: Monatsschrift für Kakteenkunde, 3:175, 1893
Cacti the coming fashion, Praha 1928
Note: Type species for the Subgenus.
schumannianus var. albispinus n.n.
schumannianus var. ampliocostatus n.n.: Note: Synonym of ampliocostatus.
schumannianus var. brevispinus n.n.
schumannianus var. nigrispinus n.n.: Note: Synonym of nigrispinus.
warasii (Ritter) Hewitt et Donald: Bradea 1:353-355, 1973
Species Catalogue for the Cactaceae, Ashingtonia 1974.

Subgenus Brasiliparodia Ritter

Note: Notobrasilia Havlicek was also a name used for this subgenus.

aurisetus n.n.
aurisetus var. longispinus n.n.
alacriportanus (Backeberg et Voll) Buxbaum: Arqu. Jard. Bot. Rio de Janeiro, 166, 1949
Die Kakteen, C VIc, Stuttgart 1. 1. 1967.
alacriportanus var. brevisetus n.n.: Note: Only a name.
alacriportanus var. cambaraensis n.n.: Note: Synonym of buenekeri var. intermedius.
alacriportanus var. fortalezensis n.n.: Note: Synonym of buenekeri fa. conjungens.
alacriportanus var. parvisetus n.n.: Note: Synonym of brevihamatus fa. conjungens.
brevihamatus (W. Haage ex Backeberg) Buxbaum [BP]: Descriptiones Cactacearum Novarum, 31, 1956
Die Kakteen, C VIc, Stuttgart 1. 1. 1967.
brevihamatus var. brasiliensis n.n.: Note: Only a name.
brevihamatus var. brunneispinus n.n.: Note: Only a name.
brevihamatus var. mollispinus (Ritter) Theunissen: Kakt. in Süd., 1:151, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
brevihamatus fa. conjungens (Ritter) Theunissen: Kakt. in Süd., 1:151, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
brevispinus n.n. [BP]: Note: Synonym of brevihamatus.
buenekeri (Buining) Buxbaum: Succulenta 41 (8):99-101, 1962
Kakt. u. a. Sukk. 17 (10):195-196, 1966
Note: Type species for the Subgenus.
buenekeri var. capeloensis n.n.: Note: buenekeri var./fa. HU 47.
buenekeri var. densispinus n.n.: Note: buenekeri var./fa. PR 263.
buenekeri var. gracilispinus n.n.: Note: buenekeri var./fa. PR 261.
buenekeri var. intermedius (Ritter) Theunissen: Kakt. in Süd., 1:151, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
buenekeri var. itambezinhoensis n.n.: Note: buenekeri FS 20.
buenekeri var. pelotasensis n.n.: Note: Synonym of catarinensis.
buenekeri var. senescens n.n.
buenekeri var. setosus n.n.: Note: Only a name.
buenekeri fa. conjungens (Ritter) Theunissen: Kakt. in Süd., 1:150, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
catarinensis (Ritter) Theunissen: Kakt. in Süd., 1:152, Spangenberg 1979
Succulenta 60 (6):140-142, 1981.
chrysocomus n.n.
chrysocomus var. elegans n.n.
chrysocomus var. rubrispinus n.n.
hamatacanthus n.n.
hamatacanthus var. albispinus n.n.
myriacanthus n.n.
myriacanthus var. longispinus n.n.
neouebelmannianus n.n.: Note: Synonym of Parodia neouebelmannianus.
parvus n.n.
rechensis Buining: Kakt. u. a. Sukk. 19 (2):23-24, 1968.

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